Anatomy of a preferred target site for the bacterial insertion sequence IS903

J Mol Biol. 2001 Feb 23;306(3):403-16. doi: 10.1006/jmbi.2000.4421.

Abstract

Like many transposons the bacterial insertion sequence IS903 was thought to insert randomly. However, using both genetic and statistical approaches, we have derived a target site for IS903 that is used 84% of the time. Computational and genetic analyses of multiple IS903 insertion sites predicted a preferred target consisting of a 21 bp palindromic pattern centered on the 9 bp target duplication generated during transposition. Here we show that targeting can be dissected into four components: the 5 bp flanking sequences, the most important sequences required for site-specific insertion; the 7 bp palindromic core within the target duplication; the dinucleotide pair at the transposon-target junction; and the local DNA context. Finally, using a substrate with multiple target sites we show that a target site is more likely found by a local bind-and-slide model and not by extended DNA tracking.

Publication types

  • Research Support, U.S. Gov't, P.H.S.

MeSH terms

  • Adenine / metabolism
  • Base Sequence
  • Computational Biology
  • Consensus Sequence / genetics
  • DNA Transposable Elements / genetics
  • DNA Transposable Elements / physiology*
  • DNA, Bacterial / genetics*
  • DNA, Bacterial / metabolism
  • Gene Duplication
  • Models, Genetic
  • Mutagenesis, Insertional / genetics
  • Plasmids / genetics
  • Recombination, Genetic / genetics*
  • Regulatory Sequences, Nucleic Acid / genetics*
  • Sequence Alignment
  • Substrate Specificity
  • Temperature

Substances

  • DNA Transposable Elements
  • DNA, Bacterial
  • Adenine