Developmental regulation of a Na(+)-activated fast outward K+ current in rat myoblasts

Cell Physiol Biochem. 2004;14(4-6):225-30. doi: 10.1159/000080331.

Abstract

Background/aims: Myoblasts undergoing differentiation sequentially express multiple K+ channels, and that ion channel expression varies depending on species and state of development. In this report, we reported a developmental regulation of fast activated and fast inactivated outward current in rat myoblasts.

Methods: The kinetic and pharmacological property of the outward current was investigated by using the patch-clamp technique.

Results: The outward current was elicited by a depolarizing step from -100 mV holding potential to +40 mV- +80 mV. The activation properties of this channel changed with days in culture. The outward current was blocked by 4-AP in a concentration dependent manner, with 0.5 mM and 2 mM 4-AP inhibiting the current by 10 +/- 3% and 56 +/- 3%, respectively. When 1 mM tetrodotoxin (TTX) was added to the bath solution or the membrane potential was depolarized to -50 mV, the fast outward current was aborted. Na+ dependent inhibition was observed when Na+ in the bath solution was replaced by Li+. In addition, replacement of K+ in the pipette solution by Cs+ almost completely eliminated the outward current.

Conclusion: The developmentally regulated outward current recorded in rat myoblasts is a Na+ influx-dependent outward K+ current, which may contribute to myoblast membrane firing of action potential or myoblast fusion.

Publication types

  • Research Support, Non-U.S. Gov't

MeSH terms

  • Animals
  • Cell Differentiation / drug effects
  • Cell Differentiation / physiology
  • Cesium / pharmacology
  • Membrane Potentials / drug effects
  • Membrane Potentials / physiology
  • Muscle Development / drug effects
  • Muscle Development / physiology*
  • Myoblasts, Skeletal / drug effects
  • Myoblasts, Skeletal / physiology*
  • Patch-Clamp Techniques
  • Potassium / pharmacology
  • Potassium / physiology
  • Potassium Channels / drug effects
  • Potassium Channels / physiology*
  • Rats
  • Sodium / pharmacology
  • Sodium / physiology
  • Sodium Channels / drug effects
  • Sodium Channels / physiology*
  • Tetrodotoxin / pharmacology

Substances

  • Potassium Channels
  • Sodium Channels
  • Cesium
  • Tetrodotoxin
  • Sodium
  • Potassium