Regulation by sphingolipids of the fate of FRTL-5 cells

J Biochem. 2009 Jan;145(1):31-6. doi: 10.1093/jb/mvn138. Epub 2008 Oct 25.

Abstract

Sphingolipids, including ceramide (Cer), sphingosine (Sph), and sphingosine 1-phosphate (Sph-1-P) have recently emerged as signal-transducing molecules. Functionally, a distinguishing characteristic of these lipids is their apparent participation in pro- or anti-proliferative cell regulation pathways. In this study, we examined the involvement of sphingolipids in the fate of FRTL-5 thyroid follicular cells. We first examined the effects of sphingolipids on FRTL-5 cell viability. Sph and Cer induced apoptosis, as revealed by fluorescence microscopy of TUNEL-positive fragmented nuclei and 180-300 bp DNA fragmentation on agarose gel electrophoresis while Sph-1-P was confirmed to prevent FRTL-5 cell apoptosis induced by deprivation of serum and TSH, possibly via cell surface receptors. We then analysed the metabolism of radiolabelled Sph and C(6)-Cer (a synthetic cell-permeable Cer) in FRTL-5 cells by thin layer chromatography, followed by autoradiography. Sph was mainly metabolized to Cer, and then to sphingomyelin, while Sph conversion into Sph-1-P was hardly detected. These changes were not affected by stimulation of the cells with TSH. Our results indicate the involvement of sphingolipid mediators in the fate of FRTL-5 thyroid cells.

MeSH terms

  • Animals
  • Cell Death
  • Cell Survival
  • Cells, Cultured
  • Ceramides / metabolism
  • Lysophospholipids / metabolism
  • Rats
  • Receptors, Cell Surface / metabolism
  • Sphingolipids / metabolism*
  • Sphingosine / analogs & derivatives
  • Sphingosine / metabolism

Substances

  • Ceramides
  • Lysophospholipids
  • Receptors, Cell Surface
  • Sphingolipids
  • sphingosine 1-phosphate
  • Sphingosine