A kinetic study of gamma-glutamyltransferase (GGT)-mediated S-nitrosoglutathione catabolism

Arch Biochem Biophys. 2009 Jan 15;481(2):191-6. doi: 10.1016/j.abb.2008.10.027. Epub 2008 Oct 29.

Abstract

S-nitrosoglutathione (GSNO) is a nitric oxide (NO) donor compound which has been postulated to be involved in transport of NO in vivo. It is known that gamma-glutamyl transpeptidase (GGT) is one of the enzymes involved in the enzyme-mediated decomposition of GSNO, but no kinetics studies of the reaction GSNO-GGT are reported in literature. In this study we directly investigated the kinetics of GGT with respect to GSNO as a substrate and glycyl-glycine (GG) as acceptor co-substrate by spectrophotometry at 334 nm. GGT hydrolyses the gamma-glutamyl moiety of GSNO to give S-nitroso-cysteinylglycine (CGNO) and gamma-glutamyl-GG. However, as both the substrate GSNO and the first product CGNO absorb at 334 nm, we optimized an ancillary reaction coupled to the enzymatic reaction, based on the copper-mediated decomposition of CGNO yielding oxidized cysteinyl-glycine and NO. The ancillary reaction allowed us to study directly the GSNO/GGT kinetics by following the decrease of the characteristic absorbance of nitrosothiols at 334 nm. A K(m) of GGT for GSNO of 0.398+/-31 mM was thus found, comparable with K(m) values reported for other gamma-glutamyl substrates of GGT.

MeSH terms

  • Animals
  • Cattle
  • Kidney / enzymology
  • Kinetics
  • Plant Proteins / metabolism
  • Plants / enzymology
  • S-Nitrosoglutathione / metabolism*
  • Solutions
  • Spectrophotometry
  • Spectrophotometry, Ultraviolet
  • Substrate Specificity
  • gamma-Glutamyltransferase / metabolism*

Substances

  • Plant Proteins
  • Solutions
  • S-Nitrosoglutathione
  • gamma-Glutamyltransferase