Fhl5/Act, a CREM-binding transcriptional activator required for normal sperm maturation and morphology, is not essential for testicular gene expression

Reprod Biol Endocrinol. 2009 Nov 24:7:133. doi: 10.1186/1477-7827-7-133.

Abstract

Background: The LIM domain protein Fhl5 was previously found to interact with CREM, a DNA binding transcriptional regulator necessary for spermiogenesis in mammals. Co-transfection experiments using heterologous promoter constructs indicated a role for Fhl5 in transcriptional up-regulation of CREM-dependent testicular genes. Male mice lacking Fhl5 were reported to be fertile but displayed partially abnormal sperm maturation and morphology.

Methods: To identify Fhl5 testicular target genes we carried out two whole-genome expression profiling experiments using high-density oligonucleotide microarrays and total testis samples from Fhl5 wild-type versus homozygous mutant mice first in different and then in isogenic strain backgrounds.

Results: Weak signal differences were detected in non-isogenic samples but no statistically significant expression changes were observed when isogenic Fhl5 mutant and wild-type samples were compared.

Conclusion: The outcome of these experiments suggests that testicular expression profiling is extremely sensitive to the genetic background and that Fhl5 is not essential for testicular gene expression to a level detected by microarray-based measurements. This might be due to redundant function of the related and similarly expressed protein Fhl4.

Publication types

  • Research Support, Non-U.S. Gov't

MeSH terms

  • Animals
  • Cyclic AMP Response Element Modulator / metabolism
  • Gene Expression Profiling
  • Gene Expression* / genetics
  • LIM Domain Proteins
  • Male
  • Mice
  • Mice, Inbred C57BL
  • Mice, Knockout
  • Oligonucleotide Array Sequence Analysis
  • Phenotype
  • Protein Binding
  • Spermatogenesis / genetics*
  • Spermatozoa / cytology*
  • Spermatozoa / metabolism
  • Spermatozoa / physiology
  • Testis / metabolism*
  • Trans-Activators / genetics
  • Trans-Activators / metabolism
  • Trans-Activators / physiology
  • Transcription Factors / genetics
  • Transcription Factors / metabolism
  • Transcription Factors / physiology*

Substances

  • Crem protein, mouse
  • Fhl5 protein, mouse
  • LIM Domain Proteins
  • Trans-Activators
  • Transcription Factors
  • Cyclic AMP Response Element Modulator