5' Triphosphorylated small interfering RNAs control replication of hepatitis B virus and induce an interferon response in human liver cells and mice

Gastroenterology. 2011 Aug;141(2):696-706, 706.e1-3. doi: 10.1053/j.gastro.2011.05.001. Epub 2011 May 13.

Abstract

Background & aims: Approved therapies for chronic hepatitis B include systemic administration of interferon (IFN)-alfa and inhibitors of hepatitis B virus (HBV) reverse-transcription. Systemic application of IFN-alfa is limited by side effects. Reverse-transcriptase inhibitors effectively control HBV replication, but rarely eliminate the virus and can select drug-resistant variants. We aimed to develop an alternative therapeutic approach that combines gene silencing with induction of IFN in the liver.

Methods: To stimulate an immune response while inhibiting HBV activity, we designed 3 small interfering (si)RNAs that target highly conserved sequences and multiple HBV transcripts of all genotypes. A 5'-triphosphate (3p) was added to the siRNAs, turning them into a ligand for the cytosolic helicase retinoic acid-inducible protein I, which becomes activated and induces expression of type-I IFNs. Antiviral activity was investigated in cell lines that replicate HBV, in HBV-infected primary human hepatocytes, and in HBV transgenic mice.

Results: 3p-double-stranded RNA (3p-RNA) activated retinoic acid-inducible protein I, induced a strong type I IFN response (expression of IFN-β) in liver cells and showed transient but strong antiviral activity. Bifunctional, HBV-specific, 3p-siRNAs controlled replication of HBV more efficiently and for longer periods of time than 3p-RNAs without silencing capacity or siRNAs that targeted identical sequences but did not contain 3p.

Conclusions: HBV-specific 3p-siRNAs are bifunctional antiviral molecules that induce production of type I IFNs in the liver and target HBV RNAs to inhibit viral replication.

Publication types

  • Research Support, Non-U.S. Gov't

MeSH terms

  • 2',5'-Oligoadenylate Synthetase / metabolism
  • Animals
  • Hep G2 Cells
  • Hepatitis B / metabolism
  • Hepatitis B / pathology
  • Hepatitis B / virology*
  • Hepatitis B virus / drug effects*
  • Hepatitis B virus / genetics*
  • Hepatitis B virus / physiology
  • Humans
  • Interferon Type I / metabolism
  • Interferon-alpha / metabolism
  • Interferon-beta / metabolism
  • Mice
  • Mice, Transgenic
  • Phosphorylation / genetics
  • RNA Interference*
  • RNA, Small Interfering / chemistry
  • RNA, Small Interfering / genetics*
  • RNA, Small Interfering / pharmacology*
  • Recombinant Proteins
  • Reverse Transcriptase Polymerase Chain Reaction
  • Trans-Activators / metabolism
  • Transcription Factors / metabolism
  • Transfection
  • Viral Load / drug effects
  • Virus Replication / drug effects*
  • Virus Replication / genetics
  • Virus Replication / physiology

Substances

  • Interferon Type I
  • Interferon-alpha
  • RAI1 protein, human
  • RNA, Small Interfering
  • Rai1 protein, mouse
  • Recombinant Proteins
  • Trans-Activators
  • Transcription Factors
  • Interferon-beta
  • 2',5'-Oligoadenylate Synthetase