Structural organization of the polysomes adjacent to mammalian processing bodies (P-bodies)

RNA Biol. 2013 Feb;10(2):314-20. doi: 10.4161/rna.23342. Epub 2013 Jan 16.

Abstract

A finely tuned balance of translation, storage and decay of mRNAs (mRNAs) is important for the regulation of gene expression. In eukaryotic cells, this takes place in dynamic cytoplasmic RNA-protein granules termed Processing bodies (P-bodies). In this study, by using immunoelectron tomography, 3D modeling and template matching, we analyze the size and the organization of the polysomes in the vicinity of human P-bodies. Our results show the presence of several polysomes that are compatible with a translational activity around P-bodies. Therefore, movement of mRNAs between polysomes and P-bodies can take place when the two compartments are in close contact. The presence of initiation factors in the proximity of P-bodies also suggests that translation of mRNAs can resume at the periphery of these granules.

Keywords: P-bodies; electron tomography; initiation; mRNA; polysomes; ribosomes.

Publication types

  • Research Support, Non-U.S. Gov't

MeSH terms

  • Cytoplasmic Granules / genetics
  • Cytoplasmic Granules / metabolism*
  • Electron Microscope Tomography
  • Eukaryotic Initiation Factor-4E / genetics
  • Eukaryotic Initiation Factor-4E / metabolism
  • Eukaryotic Initiation Factor-4G / genetics
  • Eukaryotic Initiation Factor-4G / metabolism
  • HeLa Cells
  • Humans
  • Polyribosomes / genetics
  • Polyribosomes / metabolism*
  • Polyribosomes / ultrastructure
  • Protein Biosynthesis
  • RNA Stability
  • RNA Transport*
  • RNA, Messenger / genetics
  • RNA, Messenger / metabolism

Substances

  • EIF4G2 protein, human
  • Eukaryotic Initiation Factor-4E
  • Eukaryotic Initiation Factor-4G
  • RNA, Messenger