The upstream enhancer elements of the G6PC promoter are critical for optimal G6PC expression in murine glycogen storage disease type Ia

Mol Genet Metab. 2013 Nov;110(3):275-80. doi: 10.1016/j.ymgme.2013.06.014. Epub 2013 Jun 25.

Abstract

Glycogen storage disease type-Ia (GSD-Ia) patients deficient in glucose-6-phosphatase-α (G6Pase-α or G6PC) manifest impaired glucose homeostasis characterized by fasting hypoglycemia, growth retardation, hepatomegaly, nephromegaly, hyperlipidemia, hyperuricemia, and lactic acidemia. Two efficacious recombinant adeno-associated virus pseudotype 2/8 (rAAV8) vectors expressing human G6Pase-α have been independently developed. One is a single-stranded vector containing a 2864-bp of the G6PC promoter/enhancer (rAAV8-GPE) and the other is a double-stranded vector containing a shorter 382-bp minimal G6PC promoter/enhancer (rAAV8-miGPE). To identify the best construct, a direct comparison of the rAAV8-GPE and the rAAV8-miGPE vectors was initiated to determine the best vector to take forward into clinical trials. We show that the rAAV8-GPE vector directed significantly higher levels of hepatic G6Pase-α expression, achieved greater reduction in hepatic glycogen accumulation, and led to a better toleration of fasting in GSD-Ia mice than the rAAV8-miGPE vector. Our results indicated that additional control elements in the rAAV8-GPE vector outweigh the gains from the double-stranded rAAV8-miGPE transduction efficiency, and that the rAAV8-GPE vector is the current choice for clinical translation in human GSD-Ia.

Keywords: AAV; Adeno-associated virus; G6P; G6PC promoter/enhancer; G6Pase; GPE; GSD-Ia; Gene therapy; Glucose-6-phosphatase; Glycogen storage disease type I; HCA; adeno-associated virus; glucose-6-phosphatase; glucose-6-phosphate; glycogen storage disease type Ia; hepatocellular adenoma.

Publication types

  • Research Support, N.I.H., Intramural
  • Research Support, Non-U.S. Gov't

MeSH terms

  • Animals
  • Dependovirus / genetics
  • Disease Models, Animal
  • Enhancer Elements, Genetic*
  • Gene Expression
  • Gene Expression Regulation*
  • Genetic Therapy
  • Genetic Vectors / administration & dosage
  • Genetic Vectors / genetics
  • Glucose / metabolism
  • Glucose-6-Phosphatase / genetics*
  • Glycogen Storage Disease Type I / genetics*
  • Glycogen Storage Disease Type I / metabolism
  • Humans
  • Liver / metabolism
  • Metabolome
  • Mice
  • Mice, Knockout
  • Organ Specificity
  • Promoter Regions, Genetic*
  • Transduction, Genetic
  • Transgenes

Substances

  • Glucose-6-Phosphatase
  • Glucose