Residue histidine 50 plays a key role in protecting α-synuclein from aggregation at physiological pH

J Biol Chem. 2014 May 30;289(22):15474-81. doi: 10.1074/jbc.M113.544049. Epub 2014 Apr 17.

Abstract

α-Synuclein (αSyn) aggregation is involved in the pathogenesis of Parkinson disease (PD). Recently, substitution of histidine 50 in αSyn with a glutamine, H50Q, was identified as a new familial PD mutant. Here, nuclear magnetic resonance (NMR) studies revealed that the H50Q substitution causes an increase of the flexibility of the C-terminal region. This finding provides direct evidence that this PD-causing mutant can mediate long range effects on the sampling of αSyn conformations. In vitro aggregation assays showed that substitution of His-50 with Gln, Asp, or Ala promotes αSyn aggregation, whereas substitution with the positively charged Arg suppresses αSyn aggregation. Histidine carries a partial positive charge at neutral pH, and so our result suggests that positively charged His-50 plays a role in protecting αSyn from aggregation under physiological conditions.

Keywords: Mutant; Nuclear Magnetic Resonance; Parkinson Disease; Protein Aggregation; Synuclein.

Publication types

  • Research Support, N.I.H., Extramural

MeSH terms

  • Amino Acid Substitution / physiology
  • Buffers
  • Electrochemistry
  • Histidine / metabolism*
  • Humans
  • Hydrogen-Ion Concentration
  • Nuclear Magnetic Resonance, Biomolecular
  • Parkinson Disease / genetics*
  • Parkinson Disease / metabolism*
  • Parkinson Disease / pathology
  • Unfolded Protein Response / physiology
  • alpha-Synuclein / genetics*
  • alpha-Synuclein / metabolism*

Substances

  • Buffers
  • SNCA protein, human
  • alpha-Synuclein
  • Histidine