Subcutaneous adipose tissue zinc-α2-glycoprotein is associated with adipose tissue and whole-body insulin sensitivity

Obesity (Silver Spring). 2014 Aug;22(8):1821-9. doi: 10.1002/oby.20764. Epub 2014 Apr 22.

Abstract

Objective: To examine the regulatory aspects of zinc-α2-glycoprotein (ZAG) association with obesity-related insulin resistance.

Methods: ZAG mRNA and protein were analyzed in subcutaneous adipose tissue (AT) and circulation of lean, obese, prediabetic, and type 2 diabetic men; both subcutaneous and visceral AT were explored in lean and extremely obese. Clinical and ex vivo findings were corroborated by results of in vitro ZAG silencing experiment.

Results: Subcutaneous AT ZAG was reduced in obesity, with a trend to further decrease with prediabetes and type 2 diabetes. ZAG was 3.3-fold higher in subcutaneous than in visceral AT of lean individuals. All differences were lost in extreme obesity. Obesity-associated changes in AT were not paralleled by alterations of circulating ZAG. Subcutaneous AT ZAG correlated with adiposity, adipocyte hypertrophy, whole-body and AT insulin sensitivity, mitochondrial content, expression of GLUT4, PGC1α, and adiponectin. Subcutaneous AT ZAG and adipocyte size were the only predictors of insulin sensitivity, independent on age and BMI. Silencing ZAG resulted in reduced adiponectin, IRS1, GLUT4, and PGC1α gene expression in primary human adipocytes.

Conclusions: ZAG in subcutaneous, but not in visceral AT, was markedly reduced in obesity. Clinical, cellular, and molecular evidence indicate that ZAG plays an important role in modulating whole-body and AT insulin sensitivity.

Keywords: extreme obesity; insulin sensitivity; mitochondria; subcutaneous and visceral adipose tissue; type 2 diabetes; zinc-α2-glycoprotein.

Publication types

  • Research Support, Non-U.S. Gov't

MeSH terms

  • Adipocytes / metabolism
  • Adiponectin / genetics
  • Adiponectin / metabolism
  • Adiposity / genetics
  • Adult
  • Diabetes Mellitus, Type 2 / metabolism
  • Female
  • Gene Expression
  • Gene Silencing
  • Glucose Transporter Type 4 / genetics
  • Glucose Transporter Type 4 / metabolism
  • Humans
  • Insulin Receptor Substrate Proteins / genetics
  • Insulin Receptor Substrate Proteins / metabolism
  • Insulin Resistance / genetics*
  • Intra-Abdominal Fat / metabolism*
  • Male
  • Middle Aged
  • Obesity / metabolism*
  • Peroxisome Proliferator-Activated Receptor Gamma Coactivator 1-alpha
  • Prediabetic State / metabolism
  • RNA, Messenger / metabolism
  • Seminal Plasma Proteins / genetics
  • Seminal Plasma Proteins / metabolism*
  • Subcutaneous Fat / metabolism*
  • Transcription Factors / genetics
  • Transcription Factors / metabolism
  • Zn-Alpha-2-Glycoprotein

Substances

  • Adiponectin
  • Glucose Transporter Type 4
  • IRS1 protein, human
  • Insulin Receptor Substrate Proteins
  • PPARGC1A protein, human
  • Peroxisome Proliferator-Activated Receptor Gamma Coactivator 1-alpha
  • RNA, Messenger
  • SLC2A4 protein, human
  • Seminal Plasma Proteins
  • Transcription Factors
  • Zn-Alpha-2-Glycoprotein