Abstract
Interleukin (IL)-11 has been shown to be a crucial factor for intestinal tumorigenesis, lung carcinomas, and asthma. IL-11 is thought to exclusively mediate its biological functions through cell-type-specific expression of the membrane-bound IL-11 receptor (IL-11R). Here, we show that the metalloprotease ADAM10, but not ADAM17, can release the IL-11R ectodomain. Chimeric proteins of the IL-11R and the IL-6 receptor (IL-6R) revealed that a small juxtamembrane portion is responsible for this substrate specificity of ADAM17. Furthermore, we show that the serine proteases neutrophil elastase and proteinase 3 can also cleave the IL-11R. The resulting soluble IL-11R (sIL-11R) is biologically active and binds IL-11 to activate cells. This IL-11 trans-signaling pathway can be inhibited specifically by the anti-inflammatory therapeutic compound sgp130Fc. In conclusion, proteolysis of the IL-11R represents a molecular switch that controls the IL-11 trans-signaling pathway and widens the number of cells that can be activated by IL-11.
Keywords:
ADAM10; ADAM17; interleukin-11; proteolysis; signal transduction.
Copyright © 2016 The Authors. Published by Elsevier Inc. All rights reserved.
Publication types
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Research Support, Non-U.S. Gov't
MeSH terms
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ADAM Proteins / genetics
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ADAM Proteins / immunology*
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ADAM10 Protein
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ADAM17 Protein
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Amino Acid Sequence
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Amyloid Precursor Protein Secretases / genetics
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Amyloid Precursor Protein Secretases / immunology*
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Anti-Inflammatory Agents / pharmacology
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Cell Line
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Gene Expression Regulation
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HEK293 Cells
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Humans
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Inflammation
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Interleukin-11 / genetics
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Interleukin-11 / immunology*
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Leukocyte Elastase / genetics
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Leukocyte Elastase / immunology*
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Membrane Proteins / genetics
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Membrane Proteins / immunology*
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Molecular Sequence Data
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Monocytes / drug effects
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Monocytes / immunology*
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Monocytes / pathology
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Myeloblastin / genetics
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Myeloblastin / immunology*
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Protein Binding
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Proteolysis
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Receptors, Interleukin-11 / genetics
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Receptors, Interleukin-11 / immunology*
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Receptors, Interleukin-6 / genetics
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Receptors, Interleukin-6 / immunology
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Recombinant Fusion Proteins / genetics
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Recombinant Fusion Proteins / immunology
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Recombinant Fusion Proteins / pharmacology
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Signal Transduction
Substances
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Anti-Inflammatory Agents
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IL11 protein, human
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Interleukin-11
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Membrane Proteins
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Receptors, Interleukin-11
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Receptors, Interleukin-6
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Recombinant Fusion Proteins
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Amyloid Precursor Protein Secretases
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Leukocyte Elastase
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Myeloblastin
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ADAM Proteins
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ADAM10 Protein
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ADAM10 protein, human
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ADAM17 Protein
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olamkicept