1 German Centre for Integrative Biodiversity Research (iDiv) Halle-Jena-Leipzig, Deutscher Platz 5e, 04103 Leipzig, Germany.
2 Metapopulation Research Centre, Department of Biosciences, University of Helsinki, PO Box 65, 00014 Helsinki, Finland.
3 Center for Macroecology, Evolution and Climate Change, Natural History Museum of Denmark, University of Copenhagen, Universitetsparken 15, København, DK-2100 Denmark.
4 Institute of Biology, Leipzig University, 04103 Leipzig, Germany.