Molecular characterization of transport lectin vesicular integral membrane protein 36 kDa (VIP36) in the life cycle of Schistosoma mansoni

Parasitol Res. 2017 Oct;116(10):2765-2773. doi: 10.1007/s00436-017-5587-7. Epub 2017 Aug 24.

Abstract

VIP36 is a protein described as an L-type lectin in animals, responsible for the intracellular transport of glycoproteins within the secretory pathway, and also localized on the plasma membrane. Schistosoma mansoni has a complex system of vesicles and protein transport machinery to the cell surface. The excreted/secreted products of the larvae and eggs are known to be exposed to the host immune system. Hence, characterizing the role and action of SmVIP36 in the S. mansoni life cycle is important for a better understanding of the parasite-host relationship. To this purpose, we firstly performed in silico analysis. Analysis of SmVIP36 in silico revealed that it contains a lectin leg-like domain with a jellyroll fold as seen by its putative 3D tertiary structure. Additionally, it was also observed that its CRD contains calcium ion-binding amino acids, suggesting that the binding of SmVIP36 to glycoproteins is calcium-dependent. Finally, we observed that the SmVIP36 predicted amino acid sequence relative to its orthologs was conserved. However, phylogenetic analysis revealed that SmVIP36 follows species evolution, forming a further cluster with its definitive host Homo sapiens. Moreover, q-PCR analysis in the S. mansoni life cycle points to a significant increase in gene expression in the eggs, schistosomulae, and female adult stages. Similarly, protein expression increased in eggs, cercariae, schistosomulae, and adult worm stages. These results suggest that SmVIP36 might participate in the complex secretory activity within the egg envelope and tegument proteins, both important for the stages of the parasite that interact with the host.

Keywords: L-type lectin; S. mansoni; Schistosomiasis; VIP36.

MeSH terms

  • Amino Acid Sequence
  • Animals
  • Cell Membrane / genetics
  • Cell Membrane / metabolism
  • Female
  • Gene Expression
  • Helminth Proteins / genetics*
  • Helminth Proteins / metabolism
  • Humans
  • Lectins / genetics*
  • Lectins / metabolism
  • Life Cycle Stages
  • Male
  • Membrane Proteins / genetics*
  • Membrane Proteins / metabolism
  • Mice
  • Mice, Inbred BALB C
  • Molecular Sequence Data
  • Phylogeny
  • Protein Transport
  • Schistosoma mansoni / classification
  • Schistosoma mansoni / genetics*
  • Schistosoma mansoni / growth & development*
  • Schistosoma mansoni / isolation & purification
  • Schistosomiasis mansoni / parasitology

Substances

  • Helminth Proteins
  • Lectins
  • Membrane Proteins

Grants and funding