Na+ /H+ exchanger 3 blockade ameliorates type 2 diabetes mellitus via inhibition of sodium-glucose co-transporter 1-mediated glucose absorption in the small intestine

Diabetes Obes Metab. 2018 Mar;20(3):709-717. doi: 10.1111/dom.13151. Epub 2017 Dec 5.

Abstract

Aim: To elucidate the role of Na+ /H+ exchanger 3 (NHE3) in sodium-glucose co-transporter 1 (SGLT1)-mediated small intestinal brush border membrane (BBM) glucose absorption and its functional implications in type 2 diabetes mellitus (T2DM).

Materials and methods: Human jejunal samples were obtained from patients undergoing gastrectomy. 14 C-glucose absorption was measured by liquid scintillation counting. NHE3 expression was suppressed by siRNA-mediated knockdown or augmented in Caco2 cells. Glucose and insulin tolerance in db/db and m+/db mice was assessed with oral and intraperitoneal glucose tolerance tests, and an intraperitoneal insulin tolerance test. Insulin resistance and β-cell function were assessed using homeostatic model assessment of insulin resistance and β-cell function.

Results: NHE3 expression was upregulated in db/db mouse jejunal BBM and high-glucose-treated Caco2 cells. NHE3 blockade impaired SGLT1-mediated glucose absorption in human jejunum, m+/db and db/db mouse jejunums, and Caco2 cells, via serum/glucocorticoid-regulated kinase 1 (SGK1). NHE3 knockdown suppressed SGLT1-mediated glucose uptake and reduced mRNA and protein levels of SGK1 and SGLT1, which were conversely enhanced by NHE3 overexpression. Chronic S3226 treatment diminished postprandial glucose levels and ameliorated glucose intolerance in db/db mice.

Conclusion: NHE3 is essential in the modulation of small intestinal BBM glucose absorption. Our findings provide a rationale for future possible clinical application of NHE3 for treatment of T2DM through reducing intestinal glucose uptake and counteracting postprandial hyperglycaemia.

Keywords: NHE3 inhibitor; diabetes; glucose intolerance; postprandial hyperglycaemia.

Publication types

  • Research Support, Non-U.S. Gov't

MeSH terms

  • Amiloride / analogs & derivatives
  • Amiloride / pharmacology
  • Animals
  • Caco-2 Cells
  • Diabetes Mellitus, Type 2 / drug therapy*
  • Diabetes Mellitus, Type 2 / metabolism
  • Dose-Response Relationship, Drug
  • Down-Regulation / physiology
  • Epithelial Sodium Channel Blockers / pharmacology
  • Gene Knockdown Techniques
  • Glucose / pharmacokinetics
  • Glucose Intolerance / physiopathology
  • Glucose Transporter Type 2 / metabolism
  • Humans
  • Hyperglycemia / physiopathology
  • Immediate-Early Proteins / metabolism
  • Intestinal Absorption / physiology
  • Intestinal Mucosa / metabolism
  • Intestine, Small / metabolism*
  • Male
  • Mice, Inbred C57BL
  • Postprandial Period
  • Protein Serine-Threonine Kinases / metabolism
  • Sodium-Glucose Transporter 1 / antagonists & inhibitors*
  • Sodium-Glucose Transporter 1 / metabolism
  • Sodium-Hydrogen Exchanger 3 / antagonists & inhibitors*

Substances

  • Epithelial Sodium Channel Blockers
  • Glucose Transporter Type 2
  • Immediate-Early Proteins
  • Slc2a2 protein, mouse
  • Slc5a1 protein, mouse
  • Sodium-Glucose Transporter 1
  • Sodium-Hydrogen Exchanger 3
  • Amiloride
  • Protein Serine-Threonine Kinases
  • serum-glucocorticoid regulated kinase
  • Glucose
  • ethylisopropylamiloride