Silencing of transposable elements may not be a major driver of regulatory evolution in primate iPSCs

Elife. 2018 Apr 12:7:e33084. doi: 10.7554/eLife.33084.

Abstract

Transposable elements (TEs) comprise almost half of primate genomes and their aberrant regulation can result in deleterious effects. In pluripotent stem cells, rapidly evolving KRAB-ZNF genes target TEs for silencing by H3K9me3. To investigate the evolution of TE silencing, we performed H3K9me3 ChIP-seq experiments in induced pluripotent stem cells from 10 human and 7 chimpanzee individuals. We identified four million orthologous TEs and found the SVA and ERV families to be marked most frequently by H3K9me3. We found little evidence of inter-species differences in TE silencing, with as many as 82% of putatively silenced TEs marked at similar levels in humans and chimpanzees. TEs that are preferentially silenced in one species are a similar age to those silenced in both species and are not more likely to be associated with expression divergence of nearby orthologous genes. Our data suggest limited species-specificity of TE silencing across 6 million years of primate evolution.

Keywords: Chimpanzee; Gene regulation; Transposable elements; evolution; evolutionary biology; genomics; human.

Publication types

  • Research Support, N.I.H., Extramural
  • Research Support, Non-U.S. Gov't

MeSH terms

  • Animals
  • Cells, Cultured
  • DNA Transposable Elements*
  • Epigenesis, Genetic
  • Evolution, Molecular*
  • Gene Expression Regulation*
  • Gene Silencing*
  • Genome*
  • Humans
  • Induced Pluripotent Stem Cells / cytology
  • Induced Pluripotent Stem Cells / metabolism*
  • Primates
  • Species Specificity

Substances

  • DNA Transposable Elements