1 National Park Service, Gates of the Arctic National Park & Preserve, Arctic Inventory and Monitoring Network, 4175 Geist Road, Fairbanks, Alaska, 99709, USA. kyle_joly@nps.gov.
2 Department of Biology, University of Maryland, College Park, Maryland, 20742, USA.
3 National Park Service, Yukon-Charley Rivers National Preserve, Central Alaska Inventory and Monitoring Network, 4175 Geist Road, Fairbanks, Alaska, 99709, USA.
4 Norwegian Institute for Nature Research, P.O. Box 5685, Sluppen, NO-7485, Trondheim, Norway.
5 Research Institute of Wildlife Ecology, University of Veterinary Medicine Vienna, Vienna, Austria.
6 National Park Service, Gates of the Arctic National Park & Preserve, Arctic Inventory and Monitoring Network, 4175 Geist Road, Fairbanks, Alaska, 99709, USA.
7 National Wildlife Federation, Northern Rockies, Prairies, and Pacific Region, 240 North Higgins Avenue, Suite 2, Missoula, Montana, 59802, USA.
8 National Park Service, Denali National Park and Preserve, Central Alaska Inventory and Monitoring Network, P. O. Box 9, Denali Park, Alaska, 99755, USA.
9 Senckenberg Biodiversity and Climate Research Centre, Senckenberg Gesellschaft für Naturforschung, Senckenberganlage 25, Frankfurt (Main), Germany.
10 Department of Biological Sciences, Goethe University Frankfurt, Max-von-Laue-Straße 9, 60438, Frankfurt (Main), Germany.
11 Institute of Biodiversity, Animal Health and Comparative Medicine, University of Glasgow, Glasgow, G128QQ, United Kingdom.