Rapid Chlorophyll a Fluorescence Light Response Curves Mechanistically Inform Photosynthesis Modeling

Plant Physiol. 2020 Jun;183(2):602-619. doi: 10.1104/pp.19.00375. Epub 2020 Mar 9.

Abstract

Crop improvement is crucial to ensuring global food security under climate change, and hence there is a pressing need for phenotypic observations that are both high throughput and improve mechanistic understanding of plant responses to environmental cues and limitations. In this study, chlorophyll a fluorescence light response curves and gas-exchange observations are combined to test the photosynthetic response to moderate drought in four genotypes of Brassica rapa The quantum yield of PSII (ϕ PSII ) is here analyzed as an exponential decline under changing light intensity and soil moisture. Both the maximum ϕ PSII and the rate of ϕ PSII decline across a large range of light intensities (0-1,000 μmol photons m-2 s-1; β PSII ) are negatively affected by drought. We introduce an alternative photosynthesis model (β PSII model) incorporating parameters from rapid fluorescence response curves. Specifically, the model uses β PSII as an input for estimating the photosynthetic electron transport rate, which agrees well with two existing photosynthesis models (Farquhar-von Caemmerer-Berry and Yin). The β PSII model represents a major improvement in photosynthesis modeling through the integration of high-throughput fluorescence phenotyping data, resulting in gained parameters of high mechanistic value.

Publication types

  • Research Support, Non-U.S. Gov't
  • Research Support, U.S. Gov't, Non-P.H.S.

MeSH terms

  • Brassica / metabolism*
  • Brassica / physiology*
  • Chlorophyll A / metabolism*
  • Droughts
  • Fluorescence*
  • Genotype
  • Photosynthesis / physiology
  • Photosystem II Protein Complex / metabolism

Substances

  • Photosystem II Protein Complex
  • Chlorophyll A