1 Division of Infection, Immunity & Respiratory Medicine, Faculty of Biology, Medicine and Health, School of Biological Sciences, The University of Manchester, Manchester, UK.
2 Manchester Academic Health Science Centre and NIHR Manchester Biomedical Research Unit, Manchester University NHS Foundation Trust, Manchester, UK.
3 Division of Infection, Immunity & Respiratory Medicine, Faculty of Biology, Medicine and Health, School of Biological Sciences, The University of Manchester, Manchester, UK stephen.fowler@manchester.ac.uk.