1 Key Laboratory of Utilization and Conservation for Tropical Marine Bioresources of Education of Ministry, Hainan Academy of Ocean and Fisheries Sciences, Hainan Tropical Ocean University, Haikou, China.
2 State Key Laboratory of Marine Resource Utilization in South China Sea, Hainan Aquaculture Breeding Engineering Research Center, Hainan University, Haikou, China.
3 Shenzhen Key Lab of Marine Genomics, Guangdong Provincial Key Lab of Molecular Breeding in Marine Economic Animals, BGI Academy of Marine Sciences, BGI Marine, Shenzhen, China.