Long-Term Hypoxia Maintains a State of Dedifferentiation and Enhanced Stemness in Fetal Cardiovascular Progenitor Cells

Int J Mol Sci. 2021 Aug 29;22(17):9382. doi: 10.3390/ijms22179382.

Abstract

Early-stage mammalian embryos survive within a low oxygen tension environment and develop into fully functional, healthy organisms despite this hypoxic stress. This suggests that hypoxia plays a regulative role in fetal development that influences cell mobilization, differentiation, proliferation, and survival. The long-term hypoxic environment is sustained throughout gestation. Elucidation of the mechanisms by which cardiovascular stem cells survive and thrive under hypoxic conditions would benefit cell-based therapies where stem cell survival is limited in the hypoxic environment of the infarcted heart. The current study addressed the impact of long-term hypoxia on fetal Islet-1+ cardiovascular progenitor cell clones, which were isolated from sheep housed at high altitude. The cells were then cultured in vitro in 1% oxygen and compared with control Islet-1+ cardiovascular progenitor cells maintained at 21% oxygen. RT-PCR, western blotting, flow cytometry, and migration assays evaluated adaptation to long term hypoxia in terms of survival, proliferation, and signaling. Non-canonical Wnt, Notch, AKT, HIF- and Yap1 transcripts were induced by hypoxia. The hypoxic niche environment regulates these signaling pathways to sustain the dedifferentiation and survival of fetal cardiovascular progenitor cells.

Keywords: Islet-1; cardiovascular progenitor cells; hypoxia; ovine; stemness.

MeSH terms

  • Animals
  • Cardiovascular System / cytology
  • Cardiovascular System / embryology*
  • Cell Cycle
  • Cell Differentiation
  • Cell Hypoxia / physiology*
  • Cell Movement
  • Cell Survival
  • Female
  • Hypoxia / metabolism
  • LIM-Homeodomain Proteins / genetics
  • LIM-Homeodomain Proteins / metabolism
  • Phosphatidylinositol 3-Kinases / metabolism
  • Pregnancy
  • Proto-Oncogene Proteins c-akt / metabolism
  • Sheep
  • Stem Cells / cytology*
  • Stem Cells / physiology
  • Transcription Factors / genetics
  • Transcription Factors / metabolism

Substances

  • LIM-Homeodomain Proteins
  • Transcription Factors
  • insulin gene enhancer binding protein Isl-1
  • Proto-Oncogene Proteins c-akt