The plasminogen receptor Plg-RKT regulates adipose function and metabolic homeostasis

J Thromb Haemost. 2022 Mar;20(3):742-754. doi: 10.1111/jth.15622. Epub 2021 Dec 21.

Abstract

Background: Plg-RKT , a unique transmembrane plasminogen receptor, enhances the activation of plasminogen to plasmin, and localizes the proteolytic activity of plasmin on the cell surface.

Objectives: We investigated the role of Plg-RKT in adipose function, metabolic homeostasis, and obesity.

Methods: We used adipose tissue (AT) sections from bariatric surgery patients and from high fat diet (HFD)-induced obese mice together with immunofluorescence and real-time polymerase chain reaction to study adipose expression of Plg-RKT . Mice genetically deficient in Plg-RKT and littermate controls fed a HFD or control low fat diet (LFD) were used to determine the role of Plg-RKT in insulin resistance, glucose tolerance, type 2 diabetes, and associated mechanisms including adipose inflammation, fibrosis, and ectopic lipid storage. The role of Plg-RKT in adipogenesis was determined using 3T3-L1 preadipocytes and primary cultures established from Plg-RKT -deficient and littermate control mice.

Results: Plg-RKT was highly expressed in both human and mouse AT, and its levels dramatically increased during adipogenesis. Plg-RKT -deficient mice, when fed a HFD, gained more weight, developed more hepatic steatosis, and were more insulin resistant/glucose intolerant than HFD-fed wild-type littermates. Mechanistically, these metabolic defects were linked with increased AT inflammation, AT macrophage and T-cell accumulation, adipose and hepatic fibrosis, and decreased insulin signaling in the AT and liver. Moreover, Plg-RKT regulated the expression of PPARγ and other adipogenic molecules, suggesting a novel role for Plg-RKT in the adipogenic program.

Conclusions: Plg-RKT coordinately regulates multiple aspects of adipose function that are important to maintain efficient metabolic homeostasis.

Keywords: Plg-RKT; adipose inflammation; hepatic steatosis; insulin resistance; obesity.

Publication types

  • Research Support, N.I.H., Extramural
  • Research Support, Non-U.S. Gov't
  • Research Support, U.S. Gov't, Non-P.H.S.

MeSH terms

  • Adipose Tissue* / metabolism
  • Animals
  • Diabetes Mellitus, Type 2 / metabolism
  • Dietary Fats / pharmacology
  • Fibrosis
  • Glucose Tolerance Test
  • Homeostasis*
  • Humans
  • Inflammation / metabolism
  • Insulin Resistance
  • Mice
  • Plasminogen / metabolism
  • Receptors, Cell Surface* / metabolism

Substances

  • Dietary Fats
  • PLG-R(KT) protein, mouse
  • PLGRKT protein, human
  • Receptors, Cell Surface
  • Plasminogen