PTBP1 promotes hematopoietic stem cell maintenance and red blood cell development by ensuring sufficient availability of ribosomal constituents

Cell Rep. 2022 May 10;39(6):110793. doi: 10.1016/j.celrep.2022.110793.

Abstract

Ribosomopathies constitute a range of disorders associated with defective protein synthesis mainly affecting hematopoietic stem cells (HSCs) and erythroid development. Here, we demonstrate that deletion of poly-pyrimidine-tract-binding protein 1 (PTBP1) in the hematopoietic compartment leads to the development of a ribosomopathy-like condition. Specifically, loss of PTBP1 is associated with decreases in HSC self-renewal, erythroid differentiation, and protein synthesis. Consistent with its function as a splicing regulator, PTBP1 deficiency results in splicing defects in hundreds of genes, and we demonstrate that the up-regulation of a specific isoform of CDC42 partly mimics the protein-synthesis defect associated with loss of PTBP1. Furthermore, PTBP1 deficiency is associated with a marked defect in ribosome biogenesis and a selective reduction in the translation of mRNAs encoding ribosomal proteins. Collectively, this work identifies PTBP1 as a key integrator of ribosomal functions and highlights the broad functional repertoire of RNA-binding proteins.

Keywords: CP: Molecular biology; CP: Stem cell research; PTBP1; hematopoietic stem cells; protein synthesis; red blood cell development; ribosome assembly.

Publication types

  • Research Support, Non-U.S. Gov't

MeSH terms

  • Erythrocytes / metabolism
  • Erythropoiesis
  • Hematopoietic Stem Cells* / metabolism
  • Heterogeneous-Nuclear Ribonucleoproteins / genetics
  • Heterogeneous-Nuclear Ribonucleoproteins / metabolism
  • Polypyrimidine Tract-Binding Protein / genetics
  • Polypyrimidine Tract-Binding Protein / metabolism
  • Ribosomal Proteins / metabolism
  • Ribosomes* / metabolism

Substances

  • Heterogeneous-Nuclear Ribonucleoproteins
  • Ribosomal Proteins
  • Polypyrimidine Tract-Binding Protein