1 Conservation Genetics, Beckman Center for Conservation Research, San Diego Zoo Wildlife Alliance, Escondido, CA 92027, USA.
2 San Diego Zoo Wildlife Alliance, San Diego, CA 92101, USA.
3 School of Biosciences, University of Kent, Canterbury, Kent CT2 7NJ, UK and L.K. Ernst Federal Research Center for Animal Husbandry, Dubrovitsy, Podolsk, Moscow Oblast 142132, Russia.
4 Human Genetics Department, GenoSeq Core, University of California, Los Angeles, CA 90095, USA.
5 Oregon Zoo, Portland, OR 97221, USA.
6 Disease Investigations, San Diego Zoo Wildlife Alliance, San Diego, CA 92101, USA.