ST8Sia2 polysialyltransferase protects against infection by Trypanosoma cruzi

PLoS Negl Trop Dis. 2024 Sep 25;18(9):e0012454. doi: 10.1371/journal.pntd.0012454. eCollection 2024 Sep.

Abstract

Glycosylation is one of the most structurally and functionally diverse co- and post-translational modifications in a cell. Addition and removal of glycans, especially to proteins and lipids, characterize this process which has important implications in several biological processes. In mammals, the repeated enzymatic addition of a sialic acid unit to underlying sialic acids (Sia) by polysialyltransferases, including ST8Sia2, leads to the formation of a sugar polymer called polysialic acid (polySia). The functional relevance of polySia has been extensively demonstrated in the nervous system. However, the role of polysialylation in infection is still poorly explored. Previous reports have shown that Trypanosoma cruzi (T. cruzi), a flagellated parasite that causes Chagas disease (CD), changes host sialylation of glycoproteins. To understand the role of host polySia during T. cruzi infection, we used a combination of in silico and experimental tools. We observed that T. cruzi reduces both the expression of the ST8Sia2 and the polysialylation of target substrates. We also found that chemical and genetic inhibition of host ST8Sia2 increased the parasite load in mammalian cells. We found that modulating host polysialylation may induce oxidative stress, creating a microenvironment that favors T. cruzi survival and infection. These findings suggest a novel approach to interfere with parasite infections through modulation of host polysialylation.

MeSH terms

  • Animals
  • Chagas Disease* / parasitology
  • Glycosylation
  • Humans
  • Sialic Acids* / metabolism
  • Sialyltransferases* / genetics
  • Sialyltransferases* / metabolism
  • Trypanosoma cruzi* / enzymology
  • Trypanosoma cruzi* / genetics
  • Trypanosoma cruzi* / physiology

Substances

  • Sialyltransferases
  • Sialic Acids
  • polysialic acid
  • CMP-N-acetylneuraminate-poly-alpha-2,8-sialosyl sialyltransferase

Grants and funding

We are grateful for the financial support provided by the São Paulo Research Foundation (FAPESP), grants processes n° 2018/18257-1 (GP), 2018/15549-1 (GP), 2020/04923-0 (GP), 2022/09915-0 (BRB), 2021/00140-3 (JMDS), 2022/00796-9 (LAMTS), 2021/00507-4 (VdMG), 2021/14179-9 (DMS), 2021/14751-4 (SNM), 2020/02988-7 (SKNM), 2023/02096-7 (CMM); by the Conselho Nacional de Desenvolvimento Científico e Tecnológico (“Bolsa de Produtividade”) (SKNM, CMM, and GP); by Fundação Faculdade de Medicina (FFM-SKNM); by the Coordenação de Aperfeiçoamento de Pessoal de Nível Superior (AMMN). The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript.