Investigation of Astyanax mexicanus (Characiformes, Characidae) chromosome 1 structure reveals unmapped sequences and suggests conserved evolution

PLoS One. 2024 Nov 18;19(11):e0313896. doi: 10.1371/journal.pone.0313896. eCollection 2024.

Abstract

Natural selection in the cave habitat has resulted in unique phenotypic traits (including pigmentation loss and ocular degeneration) in the Mexican tetra Astyanax mexicanus, considered a model species for evolutionary research. A. mexicanus has a karyotype of 2n = 50 chromosomes, and long-read sequencing and quantitative trait linkage maps (QTLs) have completely reconstructed the reference genome at the chromosomal level. In the current work, we performed whole chromosome isolation by microdissection and total amplification using DOP-PCR and Whole Chromosome Painting (WCP), followed by sequencing on the Illumina NextSeq platform, to investigate the microstructure of the large and conserved metacentric chromosome 1 of A. mexicanus. The sequences aligned to linkage block 3 of the reference genome, as determined by processing the reads with the DOPseq pipeline and characterizing the satellites with the TAREAN program. In addition, part of the sequences was anchored in linkage blocks that have not yet been assigned to the chromosomes. Furthermore, fluorescence in situ hybridization using WCP 1 carried out in other nearby species revealed a high degree of chromosome conservation, which allows us to hypothesize a common origin of this element. The physical mapping of the repetitive marker sequences provided a micro- and macrostructural overview and confirmed their position in chromosome pair 1. These sequences can serve as comparative tools for understanding the evolution and organization of this chromosome in other species of the family in future studies.

MeSH terms

  • Animals
  • Characidae* / genetics
  • Characiformes / genetics
  • Chromosome Mapping
  • Chromosome Painting
  • Chromosomes / genetics
  • Evolution, Molecular
  • Genetic Linkage
  • In Situ Hybridization, Fluorescence*
  • Quantitative Trait Loci

Grants and funding

This study received financial support from the Conselho Nacional de Desenvolvimento Científico e Tecnológico (CNPq) (grant 407187/2016-2). The author M.S. received a scholarship from the Conselho Nacional de Desenvolvimento Científico e Tecnológico (CNPq) (grant 160155/2018-5). The author E. H. C. de O. received a scholarship from the Conselho Nacional de Desenvolvimento Científico e Tecnológico (CNPq) (grant 307382/2019-2). https://www.gov.br/cnpq/pt-br. The author D. M. Z. A. S. received a scholarship from the Fundação de Amparo à Pesquisa do Estado de São Paulo (grant 2017/22447-8). https://fapesp.br/. Funders did not play any role in the study design, data collection and analysis, decision to publish, or preparation of the manuscript.