1 Laboratory of Wildlife Biology and Medicine, Department of Environmental Veterinary Science, Graduate School of Veterinary Medicine, Hokkaido University, Sapporo, Japan. right.rishi1@gmail.com.
2 Department of Biology, University of Kentucky, Lexington, KY, USA.
3 National Trust for Nature Conservation, Khumaltar, POB 3712, Lalitpur, Nepal.
4 Department of National Parks and Wildlife Conservation, Babarmahal, Babar Mahal, Kathmandu, Nepal.
5 Laboratory of Wildlife Biology and Medicine, Department of Environmental Veterinary Science, Graduate School of Veterinary Medicine, Hokkaido University, Sapporo, Japan.
6 One Health Research Center, Hokkaido University, Sapporo, Japan.
7 Laboratory of Wildlife Biology and Medicine, Department of Environmental Veterinary Science, Graduate School of Veterinary Medicine, Hokkaido University, Sapporo, Japan. tsubota@vetmed.hokudai.ac.jp.