Abstract
A model of the tertiary structure of human IL-6, derived from the crystal-structure of granulocyte-colony stimulating factor, reveals a 5th helical region in the loop between the first and second alpha-helix. To investigate the importance of this region for biological activity of IL-6, residues Glu-52, Ser-53, Ser-54, Lys-55, Glu-56, Leu-58, and Glu-60 were individually replaced by alanine. IL-6.Leu-58Ala displayed a 5-fold reduced biological activity on the IL-6 responsive human cell lines XG-1 and A375. This reduction in bioactivity was shown to be due to a decreased capacity of the mutant protein to trigger IL-6 receptor-alpha-chain-dependent binding to the IL-6 signal transducer, gp130.
Publication types
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Research Support, Non-U.S. Gov't
MeSH terms
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Amino Acid Sequence
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Animals
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Antigens, CD*
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Base Sequence
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Binding, Competitive
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Cell Division
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Cytokine Receptor gp130
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Humans
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Hybridomas
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Interleukin-6 / chemistry
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Interleukin-6 / genetics
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Interleukin-6 / physiology*
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Leucine / physiology*
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Membrane Glycoproteins / metabolism*
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Mice
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Models, Molecular
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Molecular Sequence Data
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Mutation
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Protein Structure, Secondary
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Protein Structure, Tertiary
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Receptors, Interleukin / metabolism
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Receptors, Interleukin-6
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Sequence Alignment
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Signal Transduction / physiology*
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Tumor Cells, Cultured
Substances
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Antigens, CD
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IL6ST protein, human
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Il6st protein, mouse
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Interleukin-6
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Membrane Glycoproteins
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Receptors, Interleukin
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Receptors, Interleukin-6
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Cytokine Receptor gp130
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Leucine